Explain with examples that Mendel’s dominant factor fails to govern phenomenon of inheritance in many cases.
By incomplete
dominance Mendel fails to govern the phenomenon of inheritance. In case of
flower colour, Mendel found that red colour of flowers was dominant over white
colour of flowers. The white coloured flowers reappeared in F2.
Later a
deviation to this rule was observed in plant species commonly called four o
clock (Mirabilis jalapa ). The plant produces flowers with red and white colours. When pure
breeding red (R) flowered plants were crossed with pure breeding white (r)
flowered plants, the F1 plants were pink (Rr) flowered. Appearance
of pink flowers, an intermediate shade between red and white, provided relief to
those who believe in blending inheritance when F2 hybrids were
crossed (Rr × Rr), the F2 generation showed a phenotypic ratio of 1
red : 2 pink : 1 white, instead of typical 3: 1 ratio. This ratio contradicted
Mendel’s principles. Appearance of red and white factors (genes) in F2
indicates that these have not been altered or blended while present together in
the pink flowered individuals. F2 reds and whites are same as the
parental red and white.
This was
declared as case of incomplete dominance. In this case neither gene is dominant
to the other. Each expresses itself in the presence of its allele to produce an
intermediate effect.
Another
important aspect of incomplete dominance in that phenotypic ratio (1: 2: 1) is
the same as genotypic ratio (1 homozygous dominant: 2 heterozygous dominant: 1
homozygous recessive) resulting from cross when two monohybrids are crossed.
When dominance is incomplete, a cross of two monohybrids (Rr × Rr) gives a
phenotypic ratio of 1: 2: 1 which is identical to the genotypic ratio. The
heterozygous individual shows incomplete dominance.
Incomplete
dominance in four o clock.
CODOMINANCE
In case of blood
types in human, both genes (A & B) produce an effect in a heterozygous
individual. This is called codominance. The genes which govern A and B blood
types are alleles. Each control the formation of a different red blood cell
Protein or antigen. Antigen a in case of person having blood group A and
antigen b in individuals with blood group B. Neither gene is dominant to the
other. The heterozygous individuals with blood group AB contains both antigen a
and b. Both proteins are detected in equal amounts in the red cells.
The case of
incomplete dominance and codominance suggest that the dominance is not
universal and absolute.
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