Describe structure and development of Anther and Microsporangia
A cross section of young anther shows that each anther lobe contains two pollen sacs or microsporangia but the number may vary among different angiosperms from one to many. As a rule microsporangia extend as longitudinally running sacs through the entire length of the anther. Later on as the anther matures, the intervening tissue breaks down between each pair of pollen sacs and the two pollen sacs coalense into one, thus forming two large cavities instead of four in each anther. In Malvaceae anthers are one called (monotherous) as there are two instead of four pollen sacs per anther and these also coalesce at maturity to form single loculus. Very rarely four celled sacs may remain separate in a mature anther. Within each pollen sac of an angiosperm as within a Microsporangium of Salaginella or Pinus are produced small numbers of micropyle mother cells surrounded by a conspicuous layer of nutritive cells, the tapetum outside the tapetum is the wall of the sporangium consistin...